-
. !- /0!12!0!
!01/"
!034!
""
0!! !00!.
1
15
6,,7#
89
:-
;&< % %= !">
# #"?#@9$%
!1
, &" < A #BC A "? ; > >BC ;? "" #BC "
D E% FGHIHJ)FGFIFJK" ?<"?
> '"! )%L< M
E BC ; #?"
A N " ?# !) t L ( # ;?
"" &O D ; < & ; HL A &" N O DE ; LF H PF HF <# &
;? Q <O ; & <O %< P <;> "LFR<O , < < ;? ; %& PR "< " FLR ,AS HR %< PR <;> " R,<O $$ET" 4
8$ )* A 0%)8 8 +" 4%$#,(? #)A <
<O;LF 7 ,
)-%?LF"?
!"
#$%&'()*+,&)
.
& &"AU V;AVS# ;A # ;A
;"?VA; ""> ;A# ; JE % J FGHIHJ) FGFIFJK A ; JJ
"? A < ; A ! U ; J) %J L < VA A
VA;E VA VU ;
AVS #"VAA;AA#!)VtL
A; < ;";AV+A
SV;HLL A" E"AA VAVLFH
PF HFW <#S +A ; VA AA "& ; < V
& < %< P U ; LFLR ;
A<+A"A ;,A ;V%&PR
" "& A A ;V &S A FLR ,AS HR %< PHR U ; R ; A , A < $$ E T" 4 8$ )*
A 0%)88 +" 4%$ A A
&,+A)A #S E;A<V;LF 7 X*/8-)# LASSF"?
" 00Y
( <" " E C HFFL A "& "
# BM D '" < ## " #
& & #BC & > < < " ; <<D#&E A< <"
< " #> & < " B ;A D & D# C &"? ; " "< ; ; ; Z & <;" & &E
;D;"? &" < A " <?< RLR6&D"
LGFG"" FGWLG "(&)PPW&"?
"<#? ;D;O;<"<E; '"#
%<$<%O%C%C%
&# && <D+ )" "?A AC* Z &O
D 1" * D &> " <'" JAJ )C " D ; #BC &O < "?#" , >"%Z&%*(PP
!"
#$%&'()*+,&)
2
(!<""DFHWA'"<< [
CD"< N #BC% ( '"# WWH A < % " ;BM # &&B";<< " D#<?)(#)&PPF
( "? O < # % C
! "< "DHWHWW A HLFPPAFR"
D ? L A PWR " D BC %*( PP
B <O # < C< ;" BC"<BC < < C"'"" D&;<C <; ?
<"D
<>BC " ;BC B" "7" <? D )C " <" < % A ! #O "? AC A '" A1 C
JQ; ?J'"O<"< < <<BC D
" <" & & " ; <" C " '" C
< < BC ; '" ! BM 6 *!I
)" PPW< R&C<#" #BC
(1D A"B" ;
< % # " A< " \*)<O<"";D;&'" #
< >;>
&;O";'"&"< >BC
#BC # " " < >BC " #( A #BC# &"<< C;" " #"C
&A &1# &" < A <BC
;?"";6"DE% 5
"
8 88 "? #O # ;? ;6
1 !
!13 0
28!8
('"! )%<!GFFH
1" A PW D > " ?< "" U <+ <" FGI)FGLFIK& "DWFHA <"<?LWU#*&
& #BC O < " &" % < "
; &"# < A " #?" &6 ( A&D O &" <? '" <? A"# (
&O AC<& " D#<?&O<O
< <O ;?"
!"
#$%&'()*+,&)
4
-/<D>BC" ?<""
'"! )%"?
& A"# C <? #& & <" O
"OWHG%<<BM # %*(PP
E " '" ! ) % D ;BC %&B '"> '" # "# "A6;# M< DPP
& ;6 ; > E " '" D
E% FGHIHJ)FGFIFJK"<#
D ; A # BC O A < # " < ; Z #BC '" "
"&&< "" #BC"
< " ?#<"&BC#">BC &
" '" & " <C 6< '" " '" < ;> #> A < [ #> D D# Q PPW
%058888
& D " ; #O O < < "
O ;" " " '" # O <& <BM "&A>" O&" ;;"
D L < M E "E?&Q<C !"
#$%&'()*+,&)
6
0$75888
5
% #?"## AN" ?#
!) t L " t < #?" '" < < " <"'"<; A'"; "
&O ; O ; "> <'" ;> < < <'"<# D " 1
758898808
+ #?" <O "C;'"
" < "BC<!;A
< ; " Q #?" #" <? " ?#"
& <? ?# ; ;" ; O &"; ""A6<"D#"
<;> " F A #?" '"
F # " " #" < " &O&"BC #?"<"N
8$
7581:
# ;? ; <O < &O D# O '" O "C C
; < &# A
D E C D " ( " A ; "? # ]&D > + < # #
"?#
& ;BC &N " #O <BM 1D
; < + < ; < 7"
< < & ; ?; & " ; ;#O"PP"^VPP
; 808
('" #;<#O+(&% ##
< ; <A )A<A < N # # "
%< #!%&
< #;?">""#&;
#; " U PH ; 6;C<O"1
#?"< ""; 6;H
; 6;""<H
( <N ;6 ; &" # N &" N # ;'_= &"
;'_= # ? # &" ? # <N ? # )A ( #?" ; &"? " N
#;EL<# !"
#$%&'()*+,&)
=
88
%< <O &A " W # > D BC ;Q
# ? < )` " "E?<))))))PP
" A<;; D"&
<;& <H!;<;" F FW WH HP & < P ; A & " ; & < W ; ;
[<;" FF A<<;" &; #<$&6& D
)$&)(# #"?#;> D
;? "'"?
&BC & B A? " +A AV ^ A P ;
# <" $& <<BC " ?< < ; < < ] ) && *"!")&*
!
0!0Y
-9
&;#< <P<O+&$& AA&D
H O "C '" ; < ; < & '" C O "C ;
8 $ A & *) < & & )%)8<$%)*%<"<A
;?&<O%< P
<;> " LFR <O '" FR ;?
< < " <O ( = '" < Q <O
;L *L L
(&#"'"? <)` C ; B
; "<#*&)L
( < & "" &A > (!)%& PPa *(&$ PPa &$%((*&( $]( PPLa **&T PPFa &*&b<( PPE<"D*(&$PP &)&" ?<%"6
&"<## <<6E[D !"
#$%&'()*+,&)
>
!
$;?<O D"E% '"! )
%"?c<OD#( Q <& Z; ]&D>+
-1.
?!3!
"1!@
/
&!&%&*&%&
8
8$+FF
$B$)A"+WHL
&!!(!&%&
A2 1"?
D#
B#
D#
"
D#
""
&"
""<
&"
%8%<+WW
<'"D<
D#
%8%%*$+WWW
<'"D& D#
1 "&
D#
+WHF
!%)8)%
%*+FFWL
B8$*+FFW
&(%\!&%&
A)
,A*K +WWL
&)+*&%&
&+WW
#
&*%&%&
""
D#
C
&"
+WHLW
(*&!&%&
8%%A+FFW
!(!&%&
<+WH
$,A!A +FF
)
%A) S+FFF
%&)&$!&%&
)8 +FW
)%
) A+WWFP
)
$B%C<+FWL
%$]S +WWFW
0%)88+"+W
A*
#$$^& $
+WWFH
A*
%E]S +WWF
)*% A])V ^ &Sc
+WW
%&*\(%&*&%&
*
K+F
%]*\)(&$&!&%&
A
^T"+WW
%$#)&%&
@&"&%AS+FFWF
%(*+&%&
D#
<"d
D#
<"d
D#
6
&"
6&
&"
< "&&"
<6
D#
;#I D#
1&D<
D#
1&D; D#
&"
<'"
D#
<"<&
D#
#
)
%,AA+WH
&
D#
8%&+FP
;"&
&"
%+FP
;"&
&"
!"
#$%&'()*+,&)
D
!
% "BC
-1.
?!3!
"1!@
/
$A+FP
;"&
A2 &"
$$ET"+FPL
A<"
D#
)
%A+FPF
& D#
%(!!&*&%&
8 A +W
$$!&%&
$+FFH
0<+F
*\+]*(E\$&%&
!*)B*<+FFH
#](*&%&
<+FP
1)%)< "&+FFPW
&&%&
<"& &"
E
#
&& A
<6
<
&"
#
B&
&"
8*
%,\U##+F
&"
8$ Ac+HWF
&"
;
D#
0
A+FH
08 A&>+>>S+FP
A%*
A+FL
<6 <6
&"
/8
)8]C+FW
""<
D#
F
*>>+WHW
<"6
D#
1
)
$,c+LFL
@
% A"U+F
*)<+F
*%8,c+F
%E A A+WWW
&+FW
$&%(#*+&%&
8$%&+
*)V>+FF
](%*&+&%&
%
E)A" +FFLH
#
D#
D#
#
#
1 D
D#
#
O<
#
>"C
D#
&& D#
<
D#
$&&%&
*A<) <+FL
#
$\+]*&%&
&<
#
$%
A+FF
&$]&%&
%&+FFF
*
$C" A+FF
*
$&<"+FFF
*
%c+FF
A%*%&<"c+FFF
%)*%<"c+FFFF
!"
<6"1
%<6
"#
&"
"<
&"
"
&"
<6
%<6
"&&"
#$%&'()*+,&)
G
!
% "C
-1.
?!3!
()&%&
"1!@
/
A2 8$8 &S+WWH
1" A
&"
)
8"U+WHH
&"
1
A+WW
H%*
%) A+WHP
%8$+WWW
)*
A+LFFL
*%)88
A+HW
(*&%&
-
#E]+FFHL
\*+&%&
%<+FFW
1"<
&"
;#&
D#
"#
D#
D#
&&C
D#
D#
"&&
D#
<
&"
1*
%+FFH
A
&"
87*+FFP
BD
#
1*
$$%c+F
(%]!&%&
<+FFL
($&%&
8+FFL
($\&$&%&
8*$8Kc+L
*#&%&
8$%*A&*A+FFPF
/88%A)A"+FFP
*,% %"+WH
*$<%A^)A+FFPL
)&!&%&
%)*8'*U+FFF
%;
,*U+FFF
1%
&)]+WH
)&(+&%&
)*%)*<c+FFWW
$*U+WH
)&*(#&%&
&)]+FFLW
,*!&%&
"&&"
<";
D#
&"
&"
#
O
#
&"
<< A
#
<"dD'" A
&"
"&
D#
<
D#
<&
D#
<?&
D#
;
#
%%8C" A+FFPP
&"
@B$C" A+F
"C
#
<+
,(%]\)&%&
<";A
48$+HH
<";A
4%$+FFW
Q
8&)]+WH
;A
D#
D#
D#
!"
#$%&'()*+,&)
I
-/.);?& ? )` D""W# WHP(!)%&PPaFW*(&$PPa
H&$%((*&($](PPLaLF**&TPPFa&*&b<(PP
&A>D ;"&#J;<# FW&*&b<(PPHa&*&b<(^&*+!)PPP BCe P($,*&PP L$()^*(&$.J&A> & <&D
*(&$PPHa()PPPa#*f(
.e&D""D%&)+*(PPFa%(!%'K(
!"
#$%&'()*+,&)
-/2);?& ? < D""W# WHP(!)%&PPaFW*(&$PPa
H&$%((*&($](PPLaLF**&TPPFa&*&b<(PP&A>
D ;"&#J;<# FW&*&b<(
PPHa&*&b<(^&*+!)PPP BCe P($,*&PP L$()^*(&$.J&A> & <&D*(&$PPHa
()PPPa#*f(
.e&D""D%&)+*(PPFa%(!%'K(
! " "< ; &A > D "&# "&"< '" < " #> &O ; "&# d
;<# %D&*&b<( PPHa&*&b<(
^ &*+!) PPP BC e ($,*& PP $() ^ *(&$ Q "? d '" "" &A > " ?<"?'" &"*PPHaPPPa#*f(
!"
#$%&'()*+,&)
.
( # '" ! ) % &# " % "? %&)+*( PPF AC %(!%'K( ; "&"<
& <O E"# # ; 8*
% ,
KU## %)* 8 ' *U * * $ &<" 8 $8 &S 8$
%&)
8 "U %;
,*U0
8 A&>+>>S-
#E]A%*
A
A%* % &<" A* # $ $ ^ & $ 1) % )< " & B8 $ * 8
7 * 8 A %
E )A" )
%A) S *,@B$C" A
-
( < < <N ;6 D " C
> +&$&
& " O #?" D ; LF ! <
PA"#" BC #?"HR= < #?"HWHR< " "
F #*& F & " O '" < &A "
"<'"D*(&$PP%&)+*(PPF"D&*&b<(
PP ($,*& PP *(&$ PPH %(!%'K( #*f(
;&$%((*&($](PPL**&TPPF&*&b<( PPH
$()^*(&$
(NO;PF& BC #?""
LPFLWR #?"!< FP
HR #?" #*& L" <<BM C < " & < " ; <?# &# '" ;
"<E**&TPPF"F
(O"N"&O'"#BC'"
<" < '" < D "> '" < ; <BC
< " ?# <>C # D " # [ '" Q '"&<O'"< #?"
" "< F ; B8 $ $$ *
&<O'"< #?" FQN;
H%*
%) $$ *
! '" > < [ ;BC #BC D ; # ; & ; < < <? A"# '" ;#
" " # &" D# # '" < <?O&; & +;HL A; ;[ # EBC=D*(&$PP#*f( &D&E ;FHFA
!"
#$%&'()*+,&)
2
!
.<O"<N;6DE% '"! )%
& O < " ?< ? # <N ,
!3!
$$
"
8
0
8?
)
0
-
-
0
0
M
M
M
.
M
@
@
HL
L
HWW
FLFL
F
LL
48$
FF
F
HP
H
HP
W
FW
PFP
)*
W
PLL
PH
HLLL
P
PH
HFW
LL
HF
0%)88
W
L
H
FP
H
WLH
L
4%$
W
PLL
FW
WWW
FL
LLHH
F
FW
F
B8$
W
W
WH
L
PFW
LH
L
H
*
$
W
LL
LLL
FW
HPP
L
LF
HLP
$
L
HHL
FLH
FFF
F
HW
HF
*
PP
LLLL
H
HH
P
PH
H%*
%)
LLL
W
FLLL
HL
WPP
LP
HWF
8$
FP
H
L
LWW
LL
HLW
L
PW
P
*%)88
L
LL
WF
WLLL
F
LPF
H
HF
F
)8
LF
W
H
W
LH
LWF
FF
H
F
A)
F
F
PH
LLL
LLH
P
LP
W
FL
1%
L
L
F
F
P
@
%
L
LLLL
L
H
FFW
L
8
8
P
F
LLL
FH
FHHP
PH
F
W
W
L
L
P
F
F
L
8
F
L
P
L
F
L
WF
A%*
L
LLL
FH
H
W
LL
HL
$%
HL
P
WW
WP
P
WP
LP
W
1*
PP
WW
WP
LL
L
L
/8
)8
W
HL
LLL
FH
H
H
LF
8%
F
WW
W
F
F
W
1
W
LLL
H
LLW
LW
H
8
W
HL
LLL
FH
HL
LF
WF
W
!%)8)%
%
W
HL
LLL
FH
P
L
W
F
8
P
F
WP
W
H
LL
WW
WW
W
F
FL
LH
%%8
W
HL
HW
L
F
F
H
LL
WW
WW
W
FFF
H
L
F
8$8
HL
P
WW
W
F
P
W
W
FH
P
PW
WP
%<
HL
P
WL
L
W
A
H
LLL
LW
FW
F
/88
P
F
WL
W
WL
PP
%
W
WL
PWH
LP
HP
%
W
LLL
H
H
F
W
%8%<
F
W
LL
WL
H
LL
P
WW
!"
#$%&'()*+,&)
4
!
.% "C
!3!
"
8
0
8?
)
0
-
-
0
0
M
M
M
. M
@
@
A*
%
F
W
LL
WL
WL
W
P
%8%%*$
HL
F
WL
H
L
H
F
8
%
F
W
LL
WL
PF
H
F
F
$B$
F
W
LL
WL
L
LH
%
H
LLL
L
P
%$
F
W
LL
WL
P
F
L
)<
LL
WW
F
PH
F
PP
W
*
$
L
WW
F
P
P
PW
L
F
W
LL
WW
F
FP
P
PF
<
L
WL
L
PL
L
!*)B*<
LL
LLL
P
FH
HW
W
*$<
)%
)
F
W
LL
WW
F
H
LH
0
L
WW
F
L
L
H
LH
0<
LL
WW
F
H
LH
*
L
WW
F
F
P
W
LL
%
L
WW
F
F
L
8
LL
WW
F
LF
F
L
L
8
LL
LLL
F
L
WH
FW
8
F
W
LL
LLL
HH
WW
F
<
H
LLL
HH
LL
WL
F
&
H
LLL
W
W
W
L
$
L
LLL
P
FH
)
8
L
LLL
FF
FH
%)*8
LL
LLL
HL
F
F
%
)
%
@
$
LL
LLL
H
L
F
LL
LLL
LH
LP
H
H
LLL
L
W
LW
F
H
LLL
FF
W
L
F
1)%
H
LLL
H
L
L
%8$
H
LLL
FH
L
F%
H
LLL
FH
L
H
LLL
LH
L
08
H
LLL
LF
L
@B$
H
LLL
P
L
P
A*
#
$
*
H
LLL
L
L
P
-
H
LLL
L
L
P
%;
H
LLL
L
L
P
)
H
LLL
L
P
!"
#$%&'()*+,&)
6
-/4&"BC Q #?"<" #;E
E% '"! )%"?
!"
-/6&"BC Q #?"<N #;EL
E% '"! )%"?
!"
#$%&'()*+,&)
=
L ;? C & P <O %< %& W ; '" &"? '"> ;?
#*& W ? # <N , %& PR
"<" " FLR ,AS HR
%< PHR & WWR <A LFR & FHR
%SFR"LFR<;> "WPR,
+&$& L & ;? '" < ; ,AS #?"%&PHP
& BC<O<"? #<N "" < < # # '" Q #?" ;
[ BC,<O
-/=?< #'"! )%"?
&<O'"&#? #<N , D"
; $$ LL <O &# N ;'_= # '" O &" ; & &"? " & <O &6 *
H %*
%) &#
# # N # < # " # N
#"
" <O " DE < [ <<"BM , + ;; W 4 8$ * 0%)88L<# !"
#$%&'()*+,&)
>
!
2?"<N;6 ? #<N E% '"! )%"?
-1.
M
%%
M
0
M
0
M
-
M
@
%&
P
W
WW
H
P
PP
W
HW
H
FL
,AS
LP
WWH
LL
HF
H
%< PP
P
PF
H
PH
&
LP
HF
FH
WW
WW
<A
LP
L
LP
WP
LF
&
WP
FLH
L
WP
FH
%S
L
PP
HH
L
F
"
W
FP
LLL
LW
LF
&<S LP
L
W
F
LWL
S
LP
LF
LL
)< LP
PH
F
FL
& W
L
PF
*"&
P
L
H
F
FF
$SA
L
P
WP
F
LF
W
HL
PL
H
P
,& W
P
LL
(<
L
HL
LF
P
L
F
F
&
L
WW
H
P
)"&
L
H
HL
]<<
L
W
LP
F
P
& A
L
P
F
SAES
L
FH
L
L
(A L
W
L
%
)<
L
LL
L
L
L
P
%"
L
H
W
L
"<A&
L
H
L
F
L
H
L
L
D ; 6; WR #?"
" ; 6;LLR; 6;LR#*&&<
; 6; D *(&$PPa&$%((*&($](
PPLa**&TPPF"&AD ;#
E< * PPa ($,*& PPa &*&b<( PPHa *(&$ PPH
& "&A
& D"##*&H '"<<L A"#
" = &>BC Q <O & # "# O & < "# &O " &>BC Q <
# '" Q<;"; <>BCD
'";&"?" AA= #BC
!"
#$%&'()*+,&)
D
!" #! !" !" -/>##BC E% '"! )%"?
$%&"
#""
-/D%"# #< BC
&"#&6'"! )%"?
(? #)A ]dLF ;"< " # > ! '" #BC " O # <## < " " #BC" !"
#$%&'()*+,&)
G
&<< A&ggg]gg
; " <;" ( D " %g g
!g'"" <;" +&$& F(Q #;Cg(< A Z <] ""<;" < '"< DD; AD<6;?<E" 7OA> &;D&;BC
<;"<;" D#< " A ; &# " ! C ] & "'"A# "BC[<<BC'"<;" " #+&$&(
# % C% ! ; " <;" ( <] D" ;
" <;" " D > ( ;
;!66;?<A> &D7 "
#< (" D '"< ;"= #BC ! < A O " <;" #;" " " #BC '" " <O " <E<< <<"<;" ; < A
!
4,D#;?'"? A D"
'"! )%"?
@2@!
&R
&; R
)R
&R
]D"
!g'7
Cg'7
%gg'7
gg'7
]g'7
&ggg'7
6;< ?#<<
%& R
!"
-"000!
I,.I
.I,4I
4I,6>
L
W
W
F
W
L
W
P
H
L
L
L
F
#$%&'()*+,&)
.I
!
6,D#;?'"? A D"
'"! )%"?
-"000!
@2@!
I,.I
.I,4I
4I,=I
&R
F
L
L
=I,G.
F
&; R
WH
H
)R
H
L
&R
W
F
&!"R
LW
]D"
L
F
W
FW
!g'7
L
Cg'7
%gg'7
F
L
gg'7
L
g
] '7
FL
LL
L
&ggg'7
W
H
6;< ?#<<
%& R
F
FL
L
WF
L
( ; "&Q Q %K&dI E A? ;#<BC"O;D" >& FL " #*& P D ; '" O <'" <" O " & <" O " # W LFG%
"O " LLq%&<"O? ";HG%# WG% HG% & <" O DE " ; LHG% #" LG% LG%&<<BC "O
#1$
$2#
$21
' ( ) *+ ) '
' *, - . /0 '
-/G%&#O& BA?" +AAV^AP<
D"E% '"! )%"?
!"
#$%&'()*+,&)
.
& <<BC <"#O ; = & FWH &% "'" "O<H <<BC <"#O < <? '" <<BC
PPL ; ;# & 1" A >&&#< <BC ""
"0!0N!-"
& #BC '" ! ) % < #
; Z ;? D " < < <
< < &D" "'""1< < < # 6< O <'" < " # & '" & #?" <'" N " ?#< <" BC
(&D#<&E< N #?"e
P " & O " D ; D "<BC
"< ?#" >BC#)A <
<O O '" < " # & " BC #<<; '" D "" #BC" ;B "<N !-!O"
/2-
&;APPL%75$9
$
8;8
P
9%8Q!BC# #
*" &"*;
&Q1$a)<,)^*<!PP%<BC;?;6
=D &"866FPW
&Q1)a)<,)a*<!^ &PPH( >BC" D
< A=D!#( e%
86DHP
&Q1)^ *PPP >BC#BC &<&%D
2F
U <<PH)S "<<A; KU*]$
$
K<" UA]"<<W
"*CPP@
%$8CV*S HF<
"*CaV%PP)$%CV*S L<
%&&<PPF%75-9
,;$
$
9,588
8+"# #
%< !"
#$%&'()*+,&)
..
%*(PP% '" Z)"?8
,!8%8
88) 1 + WL<
% BC-9
$
8;8
8
58818181)5BC# #
&"*;
)PPW%">BC )8
88
8-@
758
;?#!&+#*&7&&<
>!PPF@75$9
,
588@R
BC# #*" &"*;
"=$)a*<!^&$?;"D
#BC&"#"";6 " ?<"?'" &"".6F
*PP;875,P875
)%BP$88888%+"# #
"%< %< &)PPP-9
$
875
8
$P
9%895,
BC
# #*" &"*;
PP " ## e&7" BC<
% #BC!">%!81@-'"! )%
?W<
$<*^*<!< A"A
#BC&"#< A '"! )%<#"?
=LF
$#<" "?e+ #FH<
S!a*&a%a &aC <#S
A<; # <"FLHLHH
(#&a)<,)a%&&<^*<!PP
"D BC "?
"..L
*<!PP-
875,P5;
8
+"# #"%< %< *<!a& C,)&a)^&)PPH
< A";Q# " ?<"?'" &"
86DLW
)"<a $*a)T$ PPF* CD
! % ;= ! )D$ ;BC>?" BC'"7!#7!PPF
)))PP8
-
*-%9
#C%*(
+A AV%K^A<*P )/
% $&S;
%SDF<
!"
#$%&'()*+,&)
Download

FLORÍSTICA E FITOSSOCIOLOGIA DE UMA ÁREA DE CERRADO