!#"%$&'$#( $#( !)+*, !)- .
/103254036786 9;:<650346 =2>0@?50@AB6
CE_fmMDGFInoHJFqDGprKMRsLGDONQSXPQSutvR-SUYbTWieVXVZwbceY\gh[^cj]`_biea+]lk cedf]`ceghcjie]lk
a+_bPWPXnIxyprPyR-
+SUpTznIXYsn`_fDPM]{Rw\B|E X}s
@~l}s]`_-wbceY{a+}s]`wb_b_bkE|jceBgtWj
3ie\]`_-a+c}s]`_bke
CELpPWHNMpN `M e¡¢Fowf£¤R\FlIkmQNQBpPF[¥RXPWVQnIDSPZQS+PWYbnIXXN WIBjWQXPXBtEDOLG¦§YP¨vCk+LONM pyPXBYp SvxFq`nIeY\+nIPdY\Q e_
]lQ_bFIPWPWnII¬ªNQPWPWnIL«NQDGNQ[¥MP©pyPXgn`[¥DpXPyKªPXN}F-mMnIDOBL«p mMBF+p DGBnIFpPWBFsN PXIBNpZIF pFspmQ}F`PUP©¦UPXn`DGuN F `p[¥PHoQPXBp
°® ¯s± ²³ª´Mµ{¶\·f¸
p_fLOppx%XDPZuFlBop PFsBxyDF`DHHJDOnIKQpXLOnIDGNQPXPNMKQDFqn`x`pFNr½ BNM pXxF`©P¹ MmMPW©xpFeNIPW LGmMNMpFEPXM¹ mMDGnoPWxºHJpXNQMHDOxN IpF+nILP§IDO¦§PWxNI©P»BDO¦noFqp F+PXF`KªH¤IpFeMP¬QD¼Y
pNMn`p F-PWHJNMDDGHJ¾LGPp¿FsÀWMpePHJx¨LOmMP§LIPWÁnunI½XDGPQH½n`uDON F`H`DOxnIPHN ¿IÀWpzpy-xyn`DHJKMnIn`pXpBnIBmrPXNM¿ÀWDFqprx½Wxp JoKQPWLG¬ªPXNpXoLGDGPjF`xvpzPWKM-LGDN HmMPX`¿nIÂDG¿ÀWFspQM½XPyXxNQDGÁJH`n`DpXH¬QPjDOpLOpNMXDP NMQPXn`DPeXNMÁJIDGHPM½WDGN `nI¦XNQ¿ÀXp
zBuFqN ¦pXLG¦§PWxÃHJpxKr`ÄNrHJDPXFsKQPXnIPyPWKMnIpWÅÆmQNQMPWnsp F-HJpXNQMHDOxN IpFsNMuFloPyXn`uPM
−
−
ÉÇ È ·Ê ÈMËÌÍË
\a+cjija+kE[¥k5}swbea-]`_-c
]{]{N x`KªnIpBpXBn`ImQÏW¿NrÀWHJpDPZÎyz[¥DG¦XHpXn`LGpmQ¬M¿DOÀWppZLOpQXDGDFqP `Ðn`DHPMP[^DHJnIpX¬MDGpXLGpXDGPQ
|Bp pF+¿ÀWKMpÑnIDGNQMHJ©DGKQxPWDDHJFnI`pXDGnIKrp PWFENQDGBF`yxxpQeDGHien`pnon`PW NrPWBNMDFEF`xyXnIp mMFKªRfpxFeDGHBn`©pn`x DPWHJNMnIDpXFqnIxPXp NMF+DGHJF`xpxºpFHjÁ_fLGmMLLPXPXFIFEFqDOKMÒrn`HpBPXH¿PWÀXnIpDOÐXB`DpHFEPXFzx¡ÓDGdsHn`PXpH¤n`I ÁPWnINMDGPDF`F+xyp k+Fzno¹ _bmMPWnoPPXFIHJ£J`½rxnIDG¾DHJPXnI¿pXÀWn`p Y
PWNMDFqxp FbHpXxÔHÁLGmMLPXFsmrHPWnIDGÐW`DHPFE¡Ók+LGPF½MQmMNMXp FsznIpW`p¾p WnIDOp FI£exDHJnIpXnIPXNMDGF`xpFbPXHLGmMLGPXn`uFj¡ÆÕÖGnImQFI£J
cEnIPXNMDO¾uPX¿ÀWpvjmMLO`noPWuFlIn`mBImMnoP©Bp FsxDGHn`pn` PWNMDFqxp F\KMnIpBHPXn`DGÐWIDGHpF
hCEPWDOxKMnI`MDNQBepXFqÂHuLODLGFHmM½PXLGÅÆPXNMpnpQn` x¨hPXFsPXn`zuBPX-XnIHJmMKQLOmMPWLxPWnQNIPXpFFb¬QMPP¨H¤IHJÁÁnILODGmQPLGFPyBKMnIiepBHnIPWPXnIx DGÐW`DKrHp PQFq DO`DG¦Xpr-HJDBp F`DHJÐDGHpFhs`DGHmMnIÐXNMDHJp FhPXn`uB+HJLOmQLGPXnMPXF
¬QPPWH¤nIIÁMnIeDGPHFfLGBmMZLGPXien+noMPWPXx F-PXnINM¹ mMuPWPX`DGF¦XpQQmM[^NQ¿Â x©u¬MF-noMPWPNQP©KQPXJn`×u`Ben`DGpXHnuRLGmMLOLDGPWKrnup KrpLODFIF`PHPWnIÖBp F½ LOÖGKMDBpFszKMnIpWIÖGNQPXFj¡¢Krpn`DGNQPXF½LODGKªpXKMnIpWIÖGNQPXFo£¤
ds[^PXHJxZ`¬Mn`noDGPWÐXNrLGDGP©F`XHRhPWLGmMmMLG`PXÐXnLGDFq¡¢PXmB`pLODFqDGNQPFI£fzM`nIÐXLGDFq¡ÆLGDGF`pX¾DOx¨P £¤nIpWIpXKMLPXFq`p FfjuFlÅÆn`pKMLGPFlIpF
w\¬MnoFqPW`nINrmBPXIFsmMFqnImQPKrMPynqÒrxHDGPXx©DGFs¬MMnoPPWFsNQP¬QPXHJHJ`LOÁmQn`LGDPXPXn+FMPX+FsLO`¬QnoPPWH¤IFqÁ`nInIDGmBPIFsmMjnIP©PXnIv¹ HJmMDOu`pBPX¹ FmMÖGMxmMDGNQH¿PyÂQF+PXFsMPyxxx©x©¬MnI¬MPXnoNQPWPXNQFbP¬QHJPH¤LOImQLGnIPXDGnPXhNQhPXFpX LGDGFIFIPXHPXn`ÖBpFbMPXF-xx©Y
hw\Fq`nInIDGKMmBLGIPmMFqnIx¨PFbPQDON ت`pn`NQNrPXPXFsF-BMvPPHJBÁLOFImQÀWLGprPy ¬QPH¤`n`DPWNrPM
|mQHJLGÐDGB\c5HJnIpXxp F`F`pXx¨PZQPXFs¬QPH¤`Án`DPXFszMPF-PWno¹ mMPFLPXF`xyÖBpF
−
−
MICROBIOLOGIA
P:36191 A:2004/2005 D:1766 C:115 R:17
Página 1 de 4
!#"%$&'$#( $#( !)+*, !)- .
/103254036786 9;:<650346 =2>0@?50@AB6
w\a-PXFqDG¬ªF``pnIpmBFIFIPXIFqmMFpnIxPwPFbNQFJMÔ×Ð`F`]{KrNQn`pNQHJn`LGPmQp F-F`F MuPw\FHJF`KrÁBLOpÙmMn`LmMn`PyuLPXFq¬Q¿PXn`ÀWHJ¦§pr`PMhºn`ieDPW_bNQn`PQPWxnIDG¬rNQpUPX×B¿DÀWF`pQF`pX x¨PXFÚw\Fq`nImB`mMnoPXFÛxx©¬MnoPWNMp F`PFÑDGN `noPXHJDO`pKMLPXF`xW`DHPFÜÕuFqÖHJmMLPXF
x_bXKQN FqImQp©LGݪPyPWzXLGFqPX`nnImBIÞEmMmMnIPDGxFsDOPWpXNQIXP§LO×BpDPMPX FÖOx©¬Mn`DPXFw\Fq`nImB`mQnIPeÅÆmMNQ¿ÂFfLGPXXLOp Fhw\Fq`nImB`mQnIPQ\[^pBBLGpF-MjÝrPWLPX¿ÀXpQ[^p§¦DOY
_f_fnInIFIFIHJHJDGDGxxN N ``pypxBpDGFsHn`xp¬MDHJDGPXnIpXNMnIpyPXzNMpDGF`F`xmpFHJ pN `nIpXLGp
Ia-F`DONrnIKMDGHJPXnInINQpXpP¥NMMDGmQ¾¿PxÀXMp¥PMFq DBFlpIFzx¨xP^DHJÅÆnIupXHonIQPXPXMNMpQDGF`1x_fpmMFjnIKM¦§nIPÉpBHBPXn`DGHJÐWnIIDGFIHHJpDGFx»N `DGFIprFIàÀWp^ÞE¬MmrDGPWNQN W`nIDODÒrPMHÉP¿_fÀXDOpÉNQÁJB`pÙDHPßHJnIBpÉFIHJDGHxyn`uF`N H`DOpQxáN _fIp¥mMLO`BmQnIPâmMx¨HJpP^NKªIÖOpXNKMmQmQPQLGáP¿_fÀWp^mML¼I¬rmMPXnoH¤P Y
Mk_fPÛpNNIWDOI¬Qn`mQDOpÐXPMLO`p½ãDHJnoMpPXp»FsBHJDDGNMPXnI¿DG¬MÀWFIDHJprBDG½QxpXÒQnILON `Fsno`PXMpÑ¿PyxyÀW¬MprDDGvHJpnIFI_fpXFq¬QpÖGN DGN PX``NMunIFqpQpXzLGep»B_fp¹ pmMKªN ÖG`xynIKBpXDIHJLGDGppBpÅÆBÖGXp F`LGDGFEDGHHp»PWxNMBDprHJp nI½BF+pXMnIxPDPXFqHJNMÖGnIN DGpXF``nIxF`PWpjNQFKMeDGF`nIxkpW`N p`FDGHD+F`PMF`}½BÁKBMxyIPyDGKªHÅÆpmMFeNrnoP§¿IÀWmMpyBnoPMxF`½ªDONMKQx©ÅÆ
v¬MH¤½ªonIpUPWPX×BN NQDO`PWunsÁFNMz©DGpQMk+½ªP¨N PX`F`H¤DGÖO¬MIN DODGI¦ÐWD`F`MDeHJPXp BMF
HJDBpFbNmQHJLGDHJp F
w|×BmBDG`XnIÄDG¿NQÀWHJpyDPXxFbNDGHmBn``pnI¬MDHJDGPXDGpXNQNQP PXDGF}sDOKªpFsNmBIn`DHJDGpXNQPXDGF}noPWNrFqKªpXn``jBzNmB`nIDGN `uF
[^[^IIPXPX¬r¬rppLOLODDFqFqxxpypyxKMnIDGpHMn`pmB¬M`pDGPXn+NMBp zNMn`DGPQ
[^a-uFqIKQPXDO¬rnoPXp¿LODÀWFqpex-pyÅÆMJnIxInIpWN IIn`PÐX¿ÒrÀWHJprpÕE¡¢¹ DGmMPDGFxXDOLGpDHJn`pX LGPWÖO`NMDpXH`PXnIFÐWÒr¡Æ¦HDGp Pv£¤w xZ¬ãBN [^änIMpWÅl½¦ DPjwN `NMn CEpXmrBpXnIpWåѦ DPzKªN `p Fq ÅÆpFqÅ¢P§Ip £¤
xa-[^uFqN IKQPXoDOPX¬rno¿PXpÀX¿LODpyÀWFqp¨xHJPWpDNQB¹ PXpymMDOxynIxÐXD¬MDGFqpDIPMPXPM+mBæM`ÅÆQpXnI`nIxynIxÐWÒrN HN IpÛIPXP¿¡Ó¿¹ÀWÀWpymQpDO¬MxMmBDGpÑopXPWLGXD¼NMIHpBpWDGB`BnIpÑDGÐWpXÒªLGLçæHJH¤ÅÆp `£JDnIHJ xpræQN ÅÆIn`Px¿ÀWpyNo¬MPXmB¿IÀWÖOpnIDHPMIPWæNQÅÆÐXn`LGxDGHPMNæMoÅÆPX¿n`xÀWpNBovPXPW¿ÀWxpBDGBNMpÑpHJHJDDBBppF KMnIpXKMDGÐXNQDGHpQæMÅÆnqY
[^QcpW×B`DIp MPXF`PX¬rF`¿ÖGpNÀWLOIp©DFqxF`MEpyp©pUÅÆ×BNBpWDGI×BXppWÁPWNMÅÆnImMDGjH`pWPZvIzn`BÐXpyPXÒrNMHJÅÆpUp×BnIn`DO¡èprÅÆpXÁæ `NMpUp D×BF`HDF`PQMDON PXI¿ÁJÀWnI`ppBDHJBBpmQp¨p¿ÀWPWmÑpxÅÆpXBpX`NQzpXÖGLGPDONM`HJpWpyIn`jn`ÐXDGMP©Òrp¨HJKªpNMpX£¤D¼n+ In`
DO`PXprLOpæB¬QpU×BPXHJDGM`Pn`¿DGÀXmMp©xB p¨MDBnIDOÁNMDGpQ
[^dbDGpIFIPXFq¬rÖGN p`LODFqF`xjBpyp¬MKrDGpFIKBFq`DGN DB`ÁJpXIDGLOHDpQHPX NMpQkNIDO¬QDOÐX`DHJpFsDGNMDG¬MDBpXnIF-MPy¬MDGpFIF`ÖON `uFqzBpKªKBIDGMpXXLGDHPWNQpQ[^HPWNMDF`xyp FsBvPXH¿ÀWpr
a-ie_fpuHJNMN p¦ÁJxZIDGnoH¬MF`P©DGÀXNQpvxPXLG¿DDGHJÀWF`nIpXp¥pX¬MÁDNMPWNMDNrHÁPQP `DH_fPQ1pN§}élmMnIPXPNQ¿FlÅÆÀXppQn`x¨PLG¿PÀWFqprx1ÖB_fp pXFx KªJ`ÄNQHJDPM<}noPWNrF`Bmr¿ÀWprá}nIPXNQF`MmQ¿ÀXpÉXNMnoPWLGDG¾PXQP^F`KruHJDPWLGDO¾uPXMPQ
[^_fDDHJHJLGnIppXFenI¬MPXDGpXNMDFqxpB¹ pmMFfÖOxHJpDxyHJp p¨FPXX_fDGN HILOp^FbBXp^pPX¹ ¾mMpWÖGIxpQ¨DGHpk+F xpNMD¼ÒªHPX¿ÀWpry|D¼In`DOÒrHP¿ÀXpQc×DMP¿ÀWpPWNrPWnIÐX¬QDGP»Bp^PWxpXNMÖPXHpQa-MmQ¿ÀXp
¬MBDGuÐWF`IPDGF`HF`PDGxyBDGp¨LP§PW`¾ÐpXn``DPyprhZ|PXpBFIF`BDOmMxLPXDGLG¿PWÀX`pQÐXnIDP¨BpNMDO`noP§IpQCEF`NMDO`nID¼ÒrHPX¿ÀWpQ-DO×BP¿ÀXpBp»PX¾pWIpQDFq`x¨P¨NQD¼In`pXNrPXF`X+D¼×MP¿ÀWpF`DGx©Y
kkKMKMLOLODDHHPP¿J¿JÂÂFsFsMMP¨P¨[¥[¥DGDGHHn`n`pp¬M¬MDODOppLOLOppXXDDPPvNQPyDGNQBêQFq`nIDP©z¬QDOpX`HNMpXLGpXDGPQ½NrP¨PWXnIDHJmMLO`mMnoPM½ NMpPXxZ¬MDGN `jjNQPxMDGHDONQPQ
\a+cjija+kE[¥k@\a+}s]`_-c
S [^Á`pBBpFbjIÁHJNQDGHPXF-BvHmML¼ImMnoP
SSO ZSenInIKQF`PXNQnI¿PPy¿ÀWMpyzBxyzDmMHJnIxpXnIxyPXNMDOp¨DFqxBvpF\HmMNQLOPX`mMFsnoKªPypvDGFqnIPm»FsJKM×BLPXDFl¹ ImMN uPW`uxFsNMN p¨Ip PWn-zFqmrPyMDO¦noFqDMPXM
−
−
−
−
MICROBIOLOGIA
P:36191 A:2004/2005 D:1766 C:115 R:17
Página 2 de 4
!#"%$&'$#( $#( !)+*, !)- .
/103254036786 9;:<650346 =2>0@?50@AB6
SS ¨[^cE¬B`DGpF-NQ¿MÀXvp¨HJBmMzLO`HJmMmMnoPyLO`mMmQnoFIPXPXFsBp KQFfmMnINQPpF HJmMLO`DG¦Xp¨Bj¬QPH¤`Án`DPXF
}sÁHNMDHPXFsBvHpXLGpXnoPX¿ÀWpZBp FsxDGHn`pn` PWNMDFqxp F
MMO Sv_f_fpXpXLGLGpXpXnonoPXPX¿¿ÀWÀWp©p©MF`DGZxyieKQLOnIuPXF-x BvHJÁLOmMLPXFsxDHJnIpX¬MDPWNrPXF
MM ¨_f_fpXpXLGLGpXpXnonoPXPX¿¿ÀWÀWp©p©MMpy©ØªNQDOBMÐ L FqKª|+pXnIp©LG¬rF`PXN H¤InIDGPXNMp
MM V¨t_f_fpXpXLGLGpXpXnonoPXPX¿¿ÀWÀWp©p©QMzPyDOHNQWHKQLOmrF`mMFqLÂPyF-¬QPXMHJe`nInIDF`PWNQnI¦§P P
MTy_fpXLGpXnoPX¿ÀWp©MpyNmQHLOÐXDBj¬QPXHJ`n`DPWNMp
c5Hn`uF`HDOxNIp©xDHJnIpX¬MDPWNQp©jpF`mHJpXN In`pLOp
Q_fOpSvNÞEIn`mQpPWLOpyN IBD¼ÒrpHHJPXnI¿ÀWFIpHJDGBxp¨N HJ`nIpyFIxHJDGxyDGHn`pN ¬M`pyDPW¬QNMPXp HJ`nIDPWNMpyx%FqDFlIx¨P©ÅÆHoQPBp
EQ FqZp¨wBÒreHX`HJxDPKrMPnIPWx`mMpXnon`PX`Fse`LOÁn`¦UxPMDHPXPyFbNMxÔpHpXF`KrN `pnIn`pXp LGFfpZ¬rxPXDH¤HJInIpXnI¬MDGPXDNMPWNQpFp
kQ XykN I¦§PXF-LODPX¹ ¿mMÀWÖGxypDBHJpp FsF-PWÅÆN `DO`DGxpF+DGHDOn`NQpM¬MmMD¾PWDNMBp pF FsKªpXn+PXN `DG¬MDOÐX`DHJp FfNQpHJnIFIHJDGxyN `pBzFq`DGn`KªF-Mzw\F`HoQnIDGHoMDPyHJpXLGD
Q yw\Fq`mQBpQPyF`NrFqDG¬MDGLODMPXMvPWpFsPXNIDO¬QDOÐX`DHJpFbvF`mMLOÅ¢PWxDGQPXF PXN `DG¬MDOpXnoPWx¨PvKªpXn+BDOÅÆmQFIÀWp¨xÃxDGpFqÐLODBp
c@xIPW¬ªpXLGDFqxpyxyDHJnIpX¬QDGPXNMp
rrO ©Sew\
+FqDB`mQnIÐXBLGpDGF`MjpyJ×x`noPXIPWH¬ªLGpXmMLGLGDPXFqn-xBp¨p¨MPWPxXDGLGMmQp HpF`
rk3 yDBN F`p¨`DOÒrBHzPXmM¿xÔÀXp¨xBjDOxyp¨DF`HJnILGpXH¤nIIDOPX¦NMpDFqjxMpD¼FÅÆn`NQHJDPWL PW PWnsBzx¨PWNMDO`pLëvF`PXL
r y]lBN ID¼ÒrHPX¿ÀWpBzxDGHn`pn` PWNMDFqxp FfHpXxÃpy`uFlIvk+]-Wyw
tMt OSjk-k-H¤H¤IIDODO¦¦DDMMPXPXBMvjMHP§eoPWNML§¾`DGDx¨HP©PXF-jPXKrNIn`DOpUpU×B×BDDMM§PWIN DGIHPZFbxÔxw+w+rQHJHJpXpLGLOD D
V c5HDGHLOp¨Bp¨PX¾pWIp
tMtMO ZSew\w\FqFq``mQmQBBppQQPyP©QPXxDGBpXnIÐXNMLGDODÒrFqHzPM¿PyÀXp mMnIDP
tM ©w\Fq`mQBpQPyPF`F`DGxyDGLPX¿ÀXpyMpyNQD¼InIPW`pvMPBF`NMDO`nID¼ÒªHPX¿ÀWp
−
−
−
−
−
−
−
−
ì µl¯Eí`µl·Ê ÈMËîEË
k}-gkEã½QavQ[°ëS TBnIDGNQHJDGKMLGF-pXÅ[^DHJnIpX¬MDGpXLGpX䥡ÓWïeuBD¿ÀWp£¤ðâxf_EQdbn`p§ñ-N»mM¬MLGDGF`MnIF
kjتwÕwfCjc©½ª_ESXMhk_fÁLGmMLP©\n`pBHPXn`DGÐWIDGHPM]{N^dfDGpXLGpXDGP¨_fLGmMLPWnsv[^pLOuHJmMLPWnz¡¢XïzBD¿ÀXp £JhgDGMLç
dsgk_bò½BórriyrXXM[¥DGHn`p¬MDOpLOpXänIDONrHJDGKMLOuF+PWNrÑw×BKMLGpXnoP§IDOpNQFj¡ÓtWïzBD¿ÀXp £¤ópXNMÛðDGLOä»PWNrBpXNQF½B]{NQHW
w\a+a+w]la+kv½rð5Qbª_EQôQceEBkZ½Qórr_EQb«SXõMh[^DHJnIpX¬MDGpXLGpXDGPQÕpLÓ-ShgDGMLç
MICROBIOLOGIA
P:36191 A:2004/2005 D:1766 C:115 R:17
Página 3 de 4
!#"%$&'$#( $#( !)+*, !)- .
/103254036786 9;:<650346 =2>0@?50@AB6
g ]l[°½MCZSXõMh[^DHJnIpX¬MDGpXLGpX䥡ӧïjuBDG¿ÀWp£¤hð_bd [¥HienoPUñ
+DGLOLç
[¥nIkEN CI]qDGHij kE|v
½bPXLO[°Lã]{fN `}jn`G½bNrP§[É`DGkpXNrahPW}sLç½B]l|]{NQòZHWc© ½\ór\[°bôöëkEa+òjw\az½bóQ+SuXTdbn`pBHo÷ádbDGpXLGpXXä°pXÅv[^DGHn`ppn` PWNMDFqx¨FÛ¡¢õXï^BD¿ÀXp £¤
uBwfDG¿gÀW_-pØã£¤kE[¥azH½fie[°noPUbñ óQb
+óXnuDGLGGLÓ½-½_b]{NQ
EHXk |v½bw+-_Ebãbôøòza-]{wbiy½f|vbazSuXMù[^DHJnIpX¬QDOpLOäX3_fpNQHJKBIF»PWNQk+KQKMLODHPW`DGpXNQFÉ¡lS ï
wfa+aú©½rórMórQôÍB}kgwú©½róQQ}jSXT[^DGHn`p¬MDGpXLGpXXäCEäNQPWxDHF-PXNQCDG¦XnIF`D¼läBPWmMNrBnoF_fpXLGLOXemM¬MLGDFqMDGNMr
wfa+aú©½rórMórO½rM}kEg«wúy½QóQr}jMôÍgpXnIäÑãrWM[^DHJnIpX¬QDGPXLãgD¼ÅÆ\BPWmQNQBnoF_fpXLGLGXjmQ¬MLODF`MDONQQ
\a+wfQ_-ce}-}j½Qg\M[°O½r
kag«wúy½rórMhQôòzgw]l|v½QC©QkZrWWQh[^DHJnIpX¬MDGpXLGpXä^¡ÆXïzuBD¿ÀWp£¤hðû_bd [¥HienoPUñ
+DGLOLç
/
−
−
−
/
MICROBIOLOGIA
P:36191 A:2004/2005 D:1766 C:115 R:17
−
Página 4 de 4