??? Conorhynchos conirostris (Valenciennes, 1840) The Conorhynchos conundrum: investigating the phylogenetic position of an enigmatic neotropical catfish (Siluriformes, incertae sedis). Flávio A. Bockmann Mário C. C. de Pinna John G. Lundberg Carl J. Ferraris Jr. (FAB) Laboratório de Ictiologia de Ribeirão Preto, Dept. Biologia, FFCLRP - Universidade de São Paulo, Av. Bandeirantes 3900, 14040-901, Ribeirão Preto, SP, Brazil (MCCP) Museu de Zoologia da Universidade de São Paulo, Av. Nazaré 481, São Paulo, SP, 04263-000, Brazil (JGL) Department of Ichthyology, Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA, 19103, USA (CJF) 2944 NE Couch Street, Portland, OR, 97232, USA Conorhynchos conirostris (Valenciennes, 1840) São Francisco River Paraguassu River Conorhynchos glaber Steindachner, 1877 São Francisco River Paraguassu River Porto Seguro (Buranhaém River) Pimelodidae: Bergiaria platana Autopalatine articulation Pimelodidae: Pimelodus ornatus Conorhynchos conirostris • Fotos de vagabundagem Auchenoglanididae: Auchenoglanis occidentalis Ariidae: Arius seemanni Claroteidae: Chrysichthys sp. Pangasiidae: Pangasius sp. Conorhynchos conirostris 1) Presence of an accessory supraneural. Conorhynchos conirostris Auchenipteridae: Pseudauchenipterus nodosus Doradidae: Platydoras sp. 1) Presence of an accessory supraneural. Doradidae: Oxydoras sp. Auchenipteridae: Ageneiosus sp. Conorhynchos conirostris 2) Presence of an epioccipital process. Doradidae: Doras sp. 2) Presence of an epioccipital process. 3) Presence of a single and large foramen for the exit of the glossopharyngeal and vagus nerves. Conorhynchos conirostris Doradidae: Oxydoras sp. 4) Presence of the Müllerian ramus. Conorhynchos conirostris Doradidae: Oxydoras sp. 5) Transformator process of tripus abruptly mesially-oriented. Conorhynchos conirostris Doradidae: Doras sp. 6) Nuchal plate 3 laterally expanded. Conorhynchos conirostris Doradidae: Doras sp. 7) Mesocoracoid vestigial or absent. Conorhynchos conirostris Autapomorphies for Conorhynchos. 1) Parasphenoid and pterosphenoid largely separated from each other. Autapomorphies for Conorhynchos. 2) Deep fossa for dilatator operculi muscle. Autapomorphies for Conorhynchos. 3) Presence of the pharyngobranchial 2. ACKNOWLEDGEMENTS -National Science Foundation – “All Catfish Species Inventory” Project (NSF DEB 0315963 ). - CNPq; - FAPESP; - Aléssio Datovo and Alexandre C. Ribeiro for helping with the images.